Genus: Colonies of 4 (or 2, 8, 16) cells attached side by side, arranged linearly or zigzag; cell body elliptical or spindle or crescent in shape; terminal cells with spiny projections in many species; cell wall usually smooth, but in some species granulated or dented or ridged (Illustrations of The Japanese Fresh-water Algae, 1977). Species: Colonies of 2, 4 cells arranging linearly; cell body elongated ellipsoidal, with 1-2 short spiny projections at both ends, 9-13 μm long, 3.5-5 μm wide; each cell with longitudinal rows of small dented projections on both sides; outer cells have an additional row of dented projections arranging outside (Illustrations of The Japanese Fresh-water Algae, 1977).

50 μm 100 μm 150 μm; x 400 : 31 μm 63 μm 94 μm; x 640