|  Section [Margaritiferae]: 
Cell surface granulated, gnarled or spiny; semicells pyramidal with flattened apex or trapezoid
(Illustrations of The Japanese Fresh-water Algae, 1977). Species [f. arcticum]: 
Cell body 35 μm long, 24 μm wide, isthmus 12.6 μm wide, 
both sides with 6 waves (including basal and apical corners), apex with 2 corners and 2 waves 
(Illustrations of The Japanese Fresh-water Algae, 1977).
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